Whales, Dolphins, and Porpoises of the Western North Atlantic - Part 2
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Part 2

ORGANIZATION OF THE GUIDE

The differences between baleen and toothed whales are easy enough to see in animals washed up on the beach or maintained in a tank at a zoo or aquarium. But since an animal at sea can seldom be examined that closely, its most obvious characteristics may be its overall size, the presence or absence of a dorsal fin, its prominent coloration or markings, its general behavior, or its swimming, blowing, and diving characteristics. For that reason, regardless of their scientific relationships, all the whales, dolphins, and the one porpoise covered in the main text of this guide are divided into three groups. Those over 40 feet (12.2 m) long are discussed in the section on Large Whales, those from 13 to 40 feet (4.0 to 12.2 m) in the Medium-Sized Whale, and those less than 13 feet (4.0 m) in the Small Whales, Dolphins, and Porpoise (with a dorsal fin). There are no small whales, dolphins, or porpoises in this region without a dorsal fin. Each section is further divided into those animals with a dorsal fin and those without. From that point, animals likely to be confused in the field are grouped together and the important differences between them are discussed.

The synoptic accounts of the species are followed by five appendices: Appendix A discusses and ill.u.s.trates man-made and applied tags and natural markings on cetaceans and their importance in studies of natural history. Appendix B discusses the data which are most important to record in observations of cetaceans at sea, gives examples, and provides blank sighting forms. Appendix C discusses possible causes of cetacean strandings and the manner in which stranded animals should be handled and adds a key and tables to aid in identifying stranded cetaceans.

Appendix D provides guidelines for collecting data on stranded cetaceans and provides forms and specific instructions for taking standard measurements. Appendix E lists inst.i.tutions to be contacted in the event of a cetacean stranding or for information.

A bibliography of useful references on cetaceans in general and cetaceans of this region in particular and a directory to species accounts are included.

[Ill.u.s.tration: Figure 5.--A humpback whale lying on its left side on the deck of a Canadian whaling station. Note the fringes of baleen suspended from the roof of the mouth. (_Photo by J. G. Mead_.)]

[Ill.u.s.tration: Figure 6.--Atlantic bottlenosed dolphin mother and calf from northeastern Florida. Note the single open blowhole, a characteristic that marks these animals as toothed whales. (_Photo courtesy of Marineland of Florida._)]

[Ill.u.s.tration: Figure 7.--The open mouth of an Atlantic bottlenosed dolphin from the northeastern Gulf of Mexico. All toothed whales have teeth, which are used primarily for grasping rather than for chewing.

The number varies from 2 to over 250, though they are buried beneath the gums in females and immature animals of several species, take peculiar form in one (narwhal), and are extensively worn in others. (_Photo by D.

K. Caldwell._)]

HOW TO USE THE GUIDE

To Identify Animals at Sea

The three major sections of the guide (i.e., large, medium, and small whales) are preceded by a directory to species accounts, which is a summary of the most obvious characteristics of each species and in which summary statements about each characteristic are arranged in parallel order. To use the guide to identify living animals observed at sea, a person or persons should:

1. First estimate the animal's size and determine whether or not it has a dorsal fin.

2. Note also any distinctive features of body shape and coloration and observe its general behavior, including swimming, blowing, and diving characteristics. It should be noted that coloration may vary somewhat at sea, depending on light conditions and water clarity. For example, animals which appear dark gray or black at the surface or when dead may appear brown in good light or when submerged. Making a brief sketch at this point may aid in identifying the animal or in later recalling its distinctive features.

3. Using the directory, locate the section to which the animal probably belongs.

4. Then, for more detailed information, consult the section indicated.

There you will find a more complete discussion of the animal's range, size, and distinctive characteristics. In addition, you will find a brief discussion distinguishing it from animals with which it is likely to be confused in the field.

This guide will probably work best if, in advance of attempting to use the key in the field, the reader will familiarize himself with the general outline, with characters or behaviors to note, and with the locations of the various species accounts. It will also help if he schools himself to ask a series of questions about the animal(s) he sees at the time of the encounter rather than depending on his recall at a later time (see p. 160). As we have emphasized several times in this guide, positive identification of cetaceans at sea can only occasionally be made on the basis of a single characteristic. Therefore, the greater the amount of pertinent evidence an observer obtains, the greater the likelihood he can make a reliable identification.

To Identify Stranded Animals

Stranded animals can best be identified by referring to Appendix C and its a.s.sociated tables, making a preliminary determination and then consulting the species accounts in the main body of the book for verification of the identification. As noted in that appendix, if the animal is recently stranded, identification can be made using any of the externally visible characteristics described for the living species at sea. But even if the animal is in an advanced stage of decomposition, it can usually be identified by referring to the key and to the numbers and descriptions of baleen plates, for all baleen whales, and the numbers and relative lengths of ventral grooves, for all balaenopterine whales (Table 1), or to the tables on the numbers and descriptions of teeth, for toothed whales (Table 2).

To Record and Report Information

As discussed in the preface, though learning to identify the whales, dolphins, and porpoises one sees may be exciting in itself, many persons may want to partic.i.p.ate in the acc.u.mulation of data on these interesting animals by routinely reporting their observations to scientists who are actively studying them and who can make immediate use of the information. The following may help these persons:

Suggestions for making and recording observations of cetaceans at sea and sample data forms are included in Appendix C. Similar suggestions for taking and recording data on stranded cetaceans are included in Appendix D. For both types of data, blank data forms located after the appendices may be photocopied in bulk for use in the field.

Completed data forms and all a.s.sociated information for sightings at sea should be forwarded to the Platforms of Opportunity Program, National Marine Fisheries Service, Tiburon, CA 94920, or to one of the authors of this guide. From there, they will be made available to scientists actively studying the cetaceans of a given species or geographical area.

Completed data forms and all a.s.sociated information for observations of stranded cetaceans should be forwarded to the Division of Mammals, U.S.

National Museum, Washington, DC 20560, to one of the authors of this guide, or to one of the regional laboratories listed in Appendix E.

These persons have, in turn, been encouraged to keep a free flow of information among them.

Table 1. Ranges in Numbers of Teeth in Each Upper and Lower Jaw of Western North Atlantic Odontocetes.

[P] = Page of species account ------------------------------------------------------------------------------

Ranges in

tooth

counts

Species

-----+-----

Species common name

scientific name

[P]

Upper

Lower

Remarks --------------------+----------------+---+-----+-----+------------------------ Sperm whale

_Physeter_

57

0

18-25

Ten to sixteen upper

_catodon_

teeth _rarely_ emerge;

lower teeth fit into

sockets in upper jaw.

Northern

_Hyperoodon_

67

0

2(2)

At tip of lower jaw; bottlenosed whale

_ampullatus_

[A]

sometimes second

pair behind first.

Goosebeaked whale

_Ziphius_

70

0

2[A]

At tip of lower jaw.[B]

_cavirostris_

True's beaked whale

_Mesoplodon_

77

0

2[A]

At tip of lower jaw.[B]

_mirus_

Antillean beaked

_Mesoplodon_

78

0

2[A]

At suture of mandible.

whale

_europaeus_

One-third of way

from tip of snout to

gape.[B]

Dense-beaked whale

_Mesoplodon_

80

0

2[A]

On prominences near

_densirostris_

corner of mouth;

oriented backwards.[b]

North Sea beaked

_Mesoplodon_

82

0

2[A]

About halfway from whale

_bidens_

tip of snout to

gape.[B]

Killer whale

_Orcinus orca_

84

10-12

10-12

Prominent; curved and

oriented backwards

and inwards; pointed.

False killer whale

_Pseudorca_

88

8-11

8-11

Prominent; pointed

_cra.s.sidens_

and curved.

Atlantic pilot whale

_Globicephala_

91

8-10

8-10

-- --

_melaena_

Short-finned pilot

_Globicephala_

94

7-9

7-9

-- -- whale

_macrorhyncha_

Grampus

_Grampus_

96

0

0-7

Near front of jaw;

_griseus_

may have fallen out

in older specimens;

sometimes teeth in

upper jaw.

Beluga

_Delphinapterus_

99

8-11

8-9

-- --

_leucas_

Narwhal

_Monodon_

102

2

0[A]

One (rarely both)

_monoceros_

grows up to 9 ft

(2.5 m) tusk which

has left-hand

(sinestral) spiral.

Spotted dolphin

_Stenella_

104

30-36

28-35

-- --

_plagiodon_

Bridled dolphin

_Stenella_

108

29-34

33-36

-- --

_frontalis_

Spinner dolphin

_Stenella_

110

46-65

46-65

-- --

_longirostris_

Striped dolphin

_Stenella_

113

43-50

43-50

-- --

_coeruleoalba_

Saddleback dolphin

_Delphinus_

116

40-50

40-50

-- --

_delphis_

Fraser's dolphin

_Lagenodelphis_

120

38-44

38-44

-- --

_hosei_

Atlantic white-sided

_Lagenorhynchus_

123

30-40

30-40

Some specimens have dolphin

_acutus_

more teeth in upper

than in lower jaw.

White-beaked dolphin

_Lagenorhynchus_

126

22-28

22-28

-- --

_albirostris_

Atlantic bottlenosed

_Tursiops_

128

20-26

18-24

-- -- dolphin

_truncatus_

Guiana dolphin

_Sotalia_

132

26-35

26-35

-- --

_guianensis_

Rough-toothed

_Steno_

135

20-27

20-27

Crown is sometimes dolphin

_bredanensis_

marked with many

fine vertical

wrinkles.

Pygmy killer whale

_Feresa_

138

8-13

10-13

Many specimens have.

_attenuata_

fewer teeth on right

than on left side.

Many-toothed

_Peponocephala_

142

22-25

21-24

-- -- blackfish

_electra_

Pygmy sperm whale

_Kogia_

144

0

12-16

Rarely 10 or 11;

_breviceps_

curved back and

inwards; fit into

sockets in upper jaw.

Dwarf sperm whale

_Kogia simus_

148

0-3

8-11

Rarely 13; curved

back and in; sharply

pointed; fit into

sockets in upper jaw.

Rarely has 1-3 upper

teeth as well.

Harbor porpoise

_Phocoena_

150

22-28

22-28

Spade shaped, laterally

_phocoena_

compressed, and

relatively small.

[Footnote A: Usually erupted from gums only in adult [MALES].]

[Footnote B: May have toothpick size vestigial teeth in either jaw.]

Table 2. Body Size; Numbers, Maximum Dimensions and Descriptions of Baleen Plates; and Numbers and Relative Lengths of Ventral Grooves of W.

N. Atlantic Mysticetes.

-----------------------------------------------------------------------------

Maximum dimensions

of plates

-------------------

Number

Width

Maximum

of

Length

base

Page of

body

baleen

----+----+----+----

Species

Species

species

size[C]

plates/

ft/

ft/

common name

scientific name

account

ft(m)

side

in.

cm

in.

cm

-------------+---------------+-------+---------+--------+----+----+----+----

Blue whale

Balaenoptera

19

85 (26.0)

270-395

33"

84

12"

30

musculus

Fin whale

Balaenoptera

26

79 (24.0)

262-473

29"

72

12"

30

physalus

Sei whale

Balaenoptera

32

62 (19.0)

318-340

31"

78

15"

39

borealis

Bryde's whale

Balaenoptera

37

46 (14.0)

250-300?

17"

42

10"

24

edeni

Humpback

Megaptera

40

53 (16.0)

270-400

24"

60

5"

13

whale

novaeangliae

Bowhead whale

Balaena

49

65 (19.8)

325-360

14'

414

14"

36

mysticetus

Right whale

Eubalaena

52

53 (16.0)

250-390

7.3'

223

12"

30

glacialis

Minke whale

Balaenoptera

63

31 (10.0)

300-325

8"

21

4"

10

acutorostrata

-----------------------------------------------------------------------------

-----------------------------------------------------------------------------

Mean No.

Numbers

of

of

Species

bristles

ventral

Relative lengths common name

Color of baleen

/cm.

grooves

of ventral grooves --------------+------------------------+--------+---------+------------------ Blue whale

All black with black

10-30

55-88

At least to navel.

bristles.

Fin whale

Dark gray to bluish

10-35

56-100

At least to navel.

gray; one-fifth to

one-third of right

front is whitish.

Sei whale

Ash black with blue

35-60

38-56

End far short of

tinge and fine, light

navel.

bristles; some near

front may be light.

Bryde's whale

Slate gray with dark

15-35

40-50

At least to navel.

bristles.

Humpback whale

Ash black to olive

10-35

14-22

At least to navel.

brown; sometimes

whitish; bristles

grayish white.

Bowhead whale

Black; anterior

?

None

XX

side of some is

present.

whitish; bristles

black.

Right whale

Dirty or yellowish gray;

35-70

None

XX

some anterior plates

present.

all or part white.

Minke whale

White to yellowish

15-25

50-70

End short of

white. Posterior plates

navel; often just

may be brown or black.

behind flippers.

[Footnote C: As stated in the Index to the species, these figures represent maximum sizes recorded for the W. N. Atlantic. For all species exploited by whaling industries' current maximum sizes will be substantially smaller than these figures (see species accounts).]