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Physics tells us of the stimulus, but we are concerned with the response. The facts of color-blindness and color mixing show very clearly that the response does not tally in all respects with the stimulus. Physics, then, is apt to confuse the student at this point and lead him astray. Much impressed with the physical discovery that _white_ light is a mixture of all wave-lengths, he is ready to believe the sensation of white a mixed sensation. He says, "White is the sum of all the colors", meaning that the sensation of white is compounded of the sensations of red, orange, yellow, green, blue and violet--which is simply not true. No one can pretend to get the sensations of red or blue in the sensation of white, and the fact of complementary colors shows that you cannot tell, from the sensation of white, whether the stimulus consists of yellow and blue, or red and bluish green, or red, green and blue, or all the wave-lengths, the response being the same to all these various combinations. Total color-blindness showed us, when we were discussing this matter before, that white was an elementary sensation, and nothing that has been said since changes that conclusion.

Consider _black_, too. Physics says, black is the absence of light; but this must not be twisted to mean that black is the absence of all visual sensation. Absence of visual sensation is simply nothing, and black is far from that. It is a sensation, as positive as any, and undoubtedly elementary.

From the point of view of physics, there is no reason for considering any one color more elementary than any other. Every wave-length is elementary; and if sensation tallied precisely with the stimulus, every spectral color-tone would be an element. But there are obvious objections to such a view, such as: (1) there are not nearly as many {219} distinguishable color-tones as there are wave-lengths; (2) orange, having a single wave-length, certainly appears to be a blend as truly as purple, which has no single wave-length; and (3) we cannot get away from the fact of red-green blindness, in which there are only two color-tones, _yellow_ and _blue_. In this form of color vision (which, we must remember, is normal in the intermediate zone of the retina), there are certainly not as many elementary responses as there are wave-lengths, but only one response to all the longer waves (the sensation of yellow), one response to all the shorter waves (the sensation of blue), one response to the combination of long and short waves (the sensation of white), and one response to the cessation of light (the sensation of black). These four are certainly elementary sensations, and there are probably only a few more.

There must be at least two more, because of the fact that two of the sure elements, yellow and blue, are complementary. For suppose we try to get along with one more, as _red_. Then red, blended with yellow, would give the intervening color-tones, namely, orange with reddish and yellowish orange; and red blended with blue would give violet and purple; but yellow and blue would only give white or gray, and there would be no way of getting green. We must admit _green_ as another element. The particular red selected would be that of the red end of the spectrum, if we follow the general vote; and the green would probably be something very near gra.s.s green. We thus arrive at the conclusion that there are six elementary visual responses or sensations: white and black, yellow and blue, red and green.

It is a curious fact that some of these elementary sensations blend with each other, while some refuse to blend. White and black blend to gray, and either white or black or both together will blend with any of the four elementary colors or with any possible blend of these four. Brown, for {220} example, is a grayish orange, that is, a blend of white, black, red and yellow. Red blends with yellow, yellow with green, green with blue, and blue with red. But we cannot get yellow and blue to blend, nor red and green. When we try to get yellow and blue to blend, by combining their appropriate stimuli, both colors disappear, and we get simply the colorless sensation of white or gray.

When we try to get red and green to blend, both of them disappear and we get the sensation of yellow.

Theories of Color Vision

Of the most celebrated theories of color vision, the oldest, propounded by the physicists Young and Helmholtz, recognized only three elements, red, green and blue. Yellow they regarded as a blend of red and green, and white as a blend of all three elements. The unsatisfactory nature of this theory is obvious. White as a sensation is certainly not a blend of these three color sensations, but is, precisely, colorless; and no more is the yellow sensation a blend of red and green. Moreover, the theory cannot do justice either to total color-blindness, with its white and black but no colors, or to red-green blindness, with its yellow but no red or green.

The next prominent theory was that of the physiologist Hering. He did justice to white and black by accepting them as elements; and to yellow and blue likewise. The fact that yellow and blue would not blend he accounted for by supposing them to be antagonistic responses of the retina; when, therefore, the stimuli for both acted together on the retina, neither of the two antagonistic responses could occur, and what did occur was simply the more generic response of white.

Proceeding along this line, he concluded that red and green were also antagonistic responses; but just here {221} he committed a wholly unnecessary error, in a.s.suming that if red and green were antagonistic responses, the combination of their stimuli must give white, just as with yellow and blue. Accordingly, he was forced to select as his red and green elementary color-tones two that would be complementary; and this meant a purplish (i.e., bluish) red, and a bluish green, with the result that his "elementary" red and green appear to nearly every one as compounds and not elements. It would really have been just as easy for Hering to suppose that the red and green responses, antagonizing each other, left the sensation yellow; and then he could have selected that red and green which we have concluded above to have the best claim.

A third theory, propounded by the psychologist, Dr. Christine Ladd-Franklin, is based on keen criticism of the previous two, and seems to be harmonious with all the facts. She supposes that the color sense is now in the third stage of its evolution. In the first stage the only elements were white and black; the second stage added yellow and blue; and the third stage red and green. The outer zone of the retina is still in the first stage, and the intermediate zone in the second, only the central area having reached the third. In red-green blind individuals, the central area remains in the second stage, and in the totally color-blind the whole retina is still in the first stage.

In the first stage, one response, white, was made to light of whatever wave-length. In the second stage, this single response divided into two, one aroused by the long waves and the other by the short. The response to the long waves was the sensation of yellow, and that to the short waves the sensation of blue. In the third stage, the yellow response divided into one for the longest waves, corresponding to the red, and one for somewhat shorter waves, corresponding to the green.

Now, when we try to get a blend of red and green {222} by combining red and green lights, we fail because the two responses simply unite and revert to the more primitive yellow response; and similarly when we try to get the yellow and blue responses together, they revert to the more primitive white response out of which they developed.

But, since no one can pretend to _see_ yellow as a reddish green, nor white as a bluish yellow, it is clear that the just-spoken-of union of the red and green responses, and of the yellow and blue responses, must take place _below the level of conscious sensation_. These unions probably take place within the retina itself. Probably they are purely chemical unions.

[Ill.u.s.tration: Fig. 37--The Ladd-Franklin theory of the evolution of the color sense. (Figure text: Stage 1--white, Stage 2--yellow blue, Stage 3--red green blue)]

The _very first_ response of a rod or cone to light is probably a purely chemical reaction. Dr. Ladd-Franklin, carrying out her theory, supposes that a light-sensitive "mother substance" in the rods and cones is decomposed by the action of light, and gives off cleavage products which arouse the vital activity of the rods and cones, and thus start nerve currents coursing towards the brain.

In the "first stage", she supposes, a _single_ big cleavage product, which we may call W, is split off by the action of {223} light upon the mother substance, and the vital response to W is the sensation of white.

In the second stage, the mother substance is capable of giving off two smaller cleavage products, Y and B. Y is split off by the long waves of light, and B by the short waves, and the vital response to Y is the sensation of yellow, that to B the sensation of blue. But suppose that, chemically, Y + B = W: then, if Y and B are both split off at the same time in the same cone, they immediately unite into W, and the resulting sensation is white, and neither yellow nor blue.

[Ill.u.s.tration: Fig. 38.--The cleavage products, in the three stages of the color sense. The "mother substance" is not represented in the diagram, but only the cleavage products which, according to the Ladd-Franklin theory, are the direct stimuli for the color sensations.

(Figure text: 1--white, 2--yellow blue, 3--red green blue)]

Similarly, in the third stage, the mother substance is capable of giving off _three_ cleavage products, R, G and B; and there are three corresponding vital responses, the sensations of red, green and blue.

But, chemically, R + G = Y; and therefore, if R and G are split off at the same time, they unite chemically into Y and give the sensation of yellow. If R, G and B are all split off at the same time, they unite chemically as follows: R + G = Y, and Y + B = W; and therefore the resulting sensation is that of white.

This theory of cleavage products is in good general agreement with chemical principles, and it does justice to all the facts of color vision, as detailed in the preceding pages. It should be added that "for black, the theory supposes that, {224} in the interest of a continuous field of view, objects which reflect no light at all upon the retina have correlated with them a definite non-light sensation--that of black." [Footnote: Quotation from Dr.

Ladd-Franklin.]

Adaptation

Sensory adaptation is a change that occurs in other senses also, but it is so much more important in the sense of sight than elsewhere that it may best be considered here. The stimulus continues, the sensation ceases or diminishes--that is the most striking form of sensory adaptation. Continued action of the same stimulus puts the sense into such a condition that it responds differently from at first, and usually more weakly. It is much like fatigue, but it often is more positive and beneficial than fatigue.

The sense of smell is very subject to adaptation. On first entering a room you clearly sense an odor that you can no longer get after staying there for some time. This adaptation to one odor does not prevent your sensing quite different odors. Taste shows less adaptation than smell, but all are familiar with the decline in sweet sensation that comes with continued eating of sweets.

All of the cutaneous senses except that for pain are much subject to adaptation. Continued steady pressure gives a sensation that declines rapidly and after a time ceases altogether. The temperature sense is usually adapted to the temperature of the skin, which therefore feels neither warm nor cool. If the temperature of the skin is raised from its usual level of about 70 degrees Fahrenheit to 80 or 86, this temperature at first gives the sensation of warmth, but after a time it gives no temperature sensation at all; the warmth sense has become adapted to the temperature of 80 degrees; and now a temperature of 70 will give the sensation of cool. {225} Hold one hand in water at 80 and the other in water at 66, and when both have become adapted to these respective temperatures, plunge them together into water at 70; and you will find this last to feel cool to the warm-adapted hand and warm to the cool-adapted. There are limits to this power of adaptation.

The muscle sense seems to become adapted to any fixed position of a limb, so that, after the limb has remained motionless for some time, you cannot tell in what position it is; to find out, you have only to move it the least bit, which will excite both the muscle sense and the cutaneous pressure sense. The sense of head rotation is adaptable, in that a rotation which is keenly sensed at the start ceases to be felt as it continues; but here it is not the sense cells that become adapted, but the back flow that ceases, as will soon be explained.

To come now to the sense of sight, we have _light adaptation, dark adaptation_, and _color adaptation_. Go into a dark room, and at first all seems black, but by degrees--provided there is a little light filtering into the room--you begin to see, for your retina is becoming dark-adapted. Now go out into a bright place, and at first you are "blinded", but you quickly "get used" to the bright illumination and see objects much more distinctly than at first; for your eye has now become light-adapted. Remain for some time in a room illuminated by a colored light (as the yellowish light of most artificial illuminants), and by degrees the color sensation bleaches out so that the light appears nearly white.

Dark adaptation is equivalent to sensitizing the retina for faint light. Photographic plates can be made of more or less sensitiveness for use with different illuminations; but the retina automatically alters its sensitivity to fit the illumination to which it is exposed.

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Rod and Cone Vision

You will notice, in the dark room, that while you see light and shade and the forms of objects, you do not see colors. The same is true out of doors at night. In other words, the kind of vision that we have when the eye is dark-adapted is totally color-blind. Another significant fact is that the fovea is of little use in very dim light.

These facts are taken to mean that dim-light vision, or _twilight vision_ as it is sometimes called, is _rod vision_ and not cone vision; or, in other words, that the rods and not the cones have the great sensitiveness to faint light in the dark-adapted eye. The cones perhaps become somewhat dark-adapted, but the rods far outstrip them in this direction. The fovea has no rods and hence is of little use in very faint light. The rods have no differential responsiveness to different wave-lengths, remaining still in the "first stage" in the development of color vision, and consequently no colors are seen in faint light.

Rod vision differs then from cone vision in having only one response to every wave-length, and in adapting itself to much fainter light. No doubt, also, it is the rods that give to peripheral vision its great sensitivity to moving objects.

After-Images

After-images, which might better be called after-sensations, occur in other senses than sight, but nowhere else with such definiteness. The main fact here is that the response outlasts the stimulus. This is true of a muscle, and it is true of a sense organ. It takes a little time to get the muscle, or the sense organ, started, and, once it is in action, it takes a little time for it to stop. If you direct your eyes towards the lamp, holding your hand or a book in front of them as a screen, remove the screen for an {227} instant and then replace it, you will continue for a short time to see the light after the external stimulus has been cut off. This "positive after-image" is like the main sensation, only weaker. There is also a "negative after-image", best got by looking steadily at a black-and-white or colored figure for as long as fifteen or twenty seconds, and then directing the eyes upon a medium gray background. After a moment a sensation develops in which black takes the place of white and white of black, while for each color in the original sensation the complementary color now appears.

[Ill.u.s.tration: Fig. 39.--The visual response outlasts the stimulus.

The progress of time is supposed to be from left to right in the diagram. After the stimulus ceases, the sensation persists for a time, at first as a positive after-image, and then as a negative after-image, a sort of back swing. (Figure text: stimulus, sensory response)]

This phenomenon of the negative after-image is the same as that of color adaptation. Exposing the retina for some time to light of a certain color adapts the retina to that color, bleaches that color sensation, and, as it were, subtracts that color (or some of it) from the gray at which the eyes are then directed; and gray (or white) minus a color gives the complementary color.

Contrast

Contrast is still another effect that occurs in other senses, but most strikingly in vision. There is considerable in common between the negative after-image and contrast; indeed, {228} the negative after-image effect is also called "successive contrast". After looking at a bright surface, one of medium brightness appears dark, while this same medium brightness would seem bright after looking at a dark surface. This is evidently adaptation again, and is exactly parallel to what was found in regard to the temperature sense. After looking at any color steadily, the complementary color appears more saturated than usual; in fact, this is the way to secure the maximum of saturation in color sensation. These are examples of "successive contrast".

"Simultaneous contrast" is something new, not covered by adaptation, but gives the same effects as successive contrast. If you take two pieces of the same gray paper, and place one on a black background and the other on white, you will find the piece on the black ground to look much brighter than the piece on the white ground. Spots of gray on colored backgrounds are tinged with the complementary colors. The contrast effect is most marked at the margin adjoining the background, and grows less away from this margin. Any two adjacent surfaces produce contrast effects in each other, though we usually do not notice them any more than we usually notice the after-images that occur many times in the course of the day.

The Sense of Hearing

Sound, like light, is physically a wave motion, though the sound vibrations are very different from those of light. They travel 1,100 feet a second, instead of 186,000 miles a second. Their wave-length is measured in feet instead of in millionths of a millimeter, and their vibration frequencies are counted in tens, hundreds and thousands per second, instead of in millions of millions. But sound waves vary among themselves in the same three ways that we {229} noticed in light waves: in amplitude, in wave-length (or vibration rate), and in degree of mixture of different wave-lengths.